Molecular dynamics of MHC genesis unraveled by sequence analysis of the 1,796,938-bp HLA class I region.

نویسندگان

  • T Shiina
  • G Tamiya
  • A Oka
  • N Takishima
  • T Yamagata
  • E Kikkawa
  • K Iwata
  • M Tomizawa
  • N Okuaki
  • Y Kuwano
  • K Watanabe
  • Y Fukuzumi
  • S Itakura
  • C Sugawara
  • A Ono
  • M Yamazaki
  • H Tashiro
  • A Ando
  • T Ikemura
  • E Soeda
  • M Kimura
  • S Bahram
  • H Inoko
چکیده

The intensely studied MHC has become the paradigm for understanding the architectural evolution of vertebrate multigene families. The 4-Mb human MHC (also known as the HLA complex) encodes genes critically involved in the immune response, graft rejection, and disease susceptibility. Here we report the continuous 1,796,938-bp genomic sequence of the HLA class I region, linking genes between MICB and HLA-F. A total of 127 genes or potentially coding sequences were recognized within the analyzed sequence, establishing a high gene density of one per every 14.1 kb. The identification of 758 microsatellite provides tools for high-resolution mapping of HLA class I-associated disease genes. Most importantly, we establish that the repeated duplication and subsequent diversification of a minimal building block, MIC-HCGIX-3.8-1-P5-HCGIV-HLA class I-HCGII, engendered the present-day MHC. That the currently nonessential HLA-F and MICE genes have acted as progenitors to today's immune-competent HLA-ABC and MICA/B genes provides experimental evidence for evolution by "birth and death," which has general relevance to our understanding of the evolutionary forces driving vertebrate multigene families.

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عنوان ژورنال:
  • Proceedings of the National Academy of Sciences of the United States of America

دوره 96 23  شماره 

صفحات  -

تاریخ انتشار 1999